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19世纪中叶，达尔文发现了生物的“生命线”，创立了生物合成技术，世界的格局由此悄悄改变。德奥同盟国组成机械主义阵营，投身钢铁机甲；英法俄协约国组成达尔文主义阵营，醉心合成巨兽。1914年，世界大战爆发在即，奥匈帝国的亚历山大王子在大公遇刺后逃离祖国，身边只有几位忠臣和一台机甲为伴 。与此同时，苏格兰平民女孩德琳·夏普女扮男装加入英国空军，时时刻刻都得隐藏身份。两人以意想不到的方式登上利维坦号，少年与少女即将左右世界大战的走向…… 由橘子社Orange制作，久石让打造原创歌曲。。When three women are brutally murdered in four nights, private detective John Duval, ex-lieutenant commander in Naval Intelligence, finds himself the prime suspect in a police hunt for the sexual killer stalking Portsmouth's silent hours.。River wakes up in the mental aid department of the local hospital, not knowing how he got there. As his memory starts returning, he remembers his mission, to aid the aunt of a drug addict friend of his who committed suicide.。